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1.

Context

The development of multiple trait selection indices for solid (structure) wood production in the Scots pine (Pinus sylvestris L.) breeding program requires genetic variances and covariances estimated among wood quality traits including stiffness.

Aims

Genetic control and relationships among Scots pine growth, fiber, and wood quality traits were assessed by estimating heritability, phenotypic and genetic correlation using a Scots pine full-sib family trial.

Method

Wood quality traits including clearwood and dynamic acoustic stiffness were measured using SilviScan and Hitman in a 40-year-old progeny trial and by sampling increment cores of 778 trees of 120 families. Genetic parameters were estimated using the mixed model by the ASReml software.

Results

Heritability ranged from 0.147 to 0.306 for growth, earlywood, transition wood and latewood proportion traits and from 0.260 to 0.524 for fiber dimension, wood density, MFA and stiffness traits. The highly unfavorable genetic correlation between diameter and whole core density (?0.479) and clearwood stiffness (?0.506) and dynamic acoustic stiffness (?0.382) was observed in this study.

Conclusion

The unfavorable genetic correlations between growth traits and stiffness indicate that multiple traits selection using optimal economic weights and optimal breeding strategies are recommended for the advanced Scots pine breeding program.  相似文献   

2.

? Context

A clear understanding of the genetic control of wood properties is a prerequisite for breeding for higher wood quality in Populus tomentosa Carr. hybrid clones.

? Aims

The experiments aimed at unraveling genetic and environmental effects on wood properties among triploid hybrid clones of P. tomentosa.

? Methods

We used 5-year-old clonal trials established in Northern China to assess the heritability of wood density and fiber traits. Two hundred seventy trees from nine clones were sampled in five sites.

? Results

Site had a very significant effect on all recorded traits. Despite this large site effect, a tight genetic control was detected and clonal repeatability varied between 0.53 and 0.95. Significant genotype?×?environment interactions were detected for most of the traits. Moderate to tight correlation were evidenced among traits but they were not consistent with that in several cases that were site-dependent.

? Conclusions

Our results revealed a tight genetic control over several wood properties and therefore breeding programs might be able to improve wood density, fiber length, and coarseness in these hybrids.  相似文献   

3.

? Context

Teak??s wood color is considered an important attribute in the marketing phase and it has been influenced by environmental setting, stand conditions and management, plant genetic source, and age. However, there is a lack of understanding about how the environmental factors might affect the teak??s wood color planted in short-rotation forest plantations.

? Aims

The aim of this study is to understand the relationship, gathered from generated information, between edaphic and climatic variables and their effects in the wood color variation of Tectona grandis from trees in forest plantations.

? Methods

Twenty-two plots were grouped in five cluster sites that shared similar climatic and soil conditions. Data about soil??s physical?Cchemical properties and climatic variables were collected and analyzed. Representative trees were harvested next to each plot in order to obtain a wood sample per tree at a diameter breast height. Wood color was measured using standardized CIELab??s chromaticity system.

? Results

After comparing the wood change color index (?E*) in the five studied clusters, it was found that heartwood produced from drier and fertile sites had more yellowish-brown color. The heartwood b* color index resulted with significant correlations (R?>?0.5, P?<?0.05) among nine climatic and eight edaphic variables.

? Conclusion

It was concluded that climatic variables should be considered as the first-order causal variables to explain wood color variation. Hence, darker b* wood color was associated with dry climates; also, with deeper and fertile sites.  相似文献   

4.

? Introduction

There has been an increasing interest in very early selection of radiata pine to reduce the breeding cycle for solid wood products. For such selection, new approaches are required to assess wood quality in wood from very young stems.

? Methods

Nursery seedlings of clones of radiata pine were grown in leant condition using two leaning strategies for 18–20 months. Opposite wood and compression wood were isolated from the leaning stems and tested for dynamic modulus of elasticity, density, longitudinal shrinkage, volumetric shrinkage and compression wood area using new methods evolved for testing small size samples quickly and reliably. The methods were tested for their efficiency in differentiating clones by their wood properties.

? Results

Leaning of stems provided distinct opposite and compression wood for testing. Automated image analysis method used for compression wood area assessment was found to be a quick and effective method for processing large number of samples from young stems. Compression wood was characterised by high basic density, high longitudinal shrinkage and low volumetric shrinkage than that of opposite wood. Acoustic velocity in opposite wood had a strong negative association with longitudinal shrinkage. The study signifies the importance of preventing mixing of opposite wood with compression wood while assessing wood quality in young stems thus making leaning a critical strategy. The comparison of wood properties of opposite wood revealed significant differences between clones. Opposite wood of the clone with the lowest dynamic modulus of elasticity exhibited the highest longitudinal shrinkage.

? Conclusion

Significant differences in measurable wood properties between clones suggest the prospects of early selection for solid wood products.  相似文献   

5.

Context

Wood quality traits are important to balance the negative decline of wood quality associated with selection for growth attributes in gymnosperm breeding programs. Obtaining wood quality estimates quickly is crucial for successful incorporation in breeding programs.

Aims

The aims of this paper are to: (1) Estimate genetic and phenotypic correlations between growth and wood quality attributes, (2) Estimate heritability of the studied traits, and (3) Assess the accuracy of in situ non-destructive tools as a representative of actual wood density.

Methods

Wood density (X-ray densitometry), tree height, diameter, volume, resistance drilling, acoustic velocity, and dynamic modulus of elasticity were estimated, along with their genetic parameters, for 1,200, 20-year-old trees from 25 open-pollinated families.

Results

Individual tree level heritabilities for non-destructive evaluation attributes were moderate ( $ {\widehat{h}}_i^2=0.37-0.42 $ ), wood density and growth traits were lower ( $ {\widehat{h}}_i^2=0.23-0.35 $ ). Favorable genetic and phenotypic correlations between growth traits, wood density, and non-destructive evaluation traits were observed. A perfect genetic correlation was found between resistance drilling and wood density (r G ?=?1.00?±?0.07), while acoustic velocity and dynamic modulus of elasticity showed weaker genetic correlations with wood density (r G ?=?0.25?±?0.24;?0.46?±?0.21, respectively).

Conclusion

This study confirmed that resistance drilling is a reliable predictor of wood density in western larch, while the weak genetic correlations displayed by acoustic velocity and dynamic modulus of elasticity suggest limited dependability for their use as fast in situ wood density assessment methods in this species.  相似文献   

6.

Context

The knowledge of consumer preferences in wood furniture is crucial for the wood processing industries.

Aim

This study aims to identify the attributes of wood that affect the preferences of consumers from the city of Antananarivo, Madagascar, for wooden furnishings.

Methods

Quantitative measurements of the density and aesthetic properties of 12 wood species were carried out. The properties measured were the colour in the CIEL*a*b* colour space, the texture through greyscale image processing, and the density. Then, the wood specimens were subjected to sensory analysis with 100 consumers.

Results

The results showed that the "density" was a prime criterion in choosing a wood species. Concerning the visual aspect, Antananarivo consumers generally liked slightly dark wood colour (L* around 52), tending towards yellow, with a visible oriented texture. The influence of socio-economic factors was also highlighted.

Conclusion

These results are of prime importance in the search for alternative species to replace precious woods that are becoming rare.  相似文献   

7.

? Context

The correlation between tree ring width and density and short-term climate fluctuations may be a useful tool for predicting response of wood formation process to long-term climate change.

? Aims

This study examined these correlations for different radiata pine genotypes and aimed at detecting potential genotype by climate interactions.

? Methods

Four data sets comprising ring width and density of half- and full-sib radiata pine families were used. Correlations with climate variables were examined, after the extraction of the effect of cambial age.

? Results

Cambial age explained the highest proportion of the ring to ring variation in all variables. Calendar year and year by family interaction explained a smaller but significant proportion of the variation. Rainfall had a positive correlation with ring width and, depending on test site, either a negative or positive correlation with ring density. Correlations between temperature during growing season and ring density were generally negative.

? Conclusion

Climate variables that influence ring width and wood density can be identified from ring profiles, after removing the cambial age effect. Families can be selected that consistently show desirable response to climate features expected to become prevalent as a result of climate change.  相似文献   

8.
9.

Context

Reliable estimates of wood density (WD) within individual trees could maximize the value of Pinus massoniana for specific end-use.

Aim

We examined and quantified the axial patterns of WD in trees with different social status in the stands.

Methods

Wood disks were sampled at the bottom, breast height, and middle of each 1-m sections from 108 stems, harvesting from three social classes in subtropical forests. A mixed-effects model was designed to quantify axial variation.

Results

The WD at different height was significantly different from the whole-stem WD (WWD) except the relative height of 0.1. An overall decrease of 133.8 kg m?3 in WD was found from stem base to top. WD was significantly influenced by relative heights, tree age, and social class. WD of each relative height in mature trees was significantly higher than that of younger trees. Tree social class can affect WD development in the axial direction at age classes 2 and 3. Combining the fixed plus random effects, the final model explained 91 % of the observed variation in WD.

Conclusion

The WD development patterns in the axial direction vary considerably among tree age, diameter at breast height, and social class. To distinguish and supply timber for specific end-uses, we should use the axial variation in disk WD (DWD) instead of WWD directly. The accurate predictions of WD provided by the model could be used to optimally classify logs into different product classes and maximize economic benefits. We can use DWD at the relative height of 0.1 instead of WWD of a single tree.  相似文献   

10.

Aims

The objective of this study was to compare the merit of the Colombian landrace relative to the various Australian native races of Eucalyptus globulus Labill. ssp. globulus and study the genetic control of key traits such as growth, wood density, and leaf phase change in the unique conditions of the Colombian highlands.

Methods

The genetic study was based on open-pollinated families from native Australian and Colombian landrace origin, tested across four trials spanning two generations of breeding. A multisite mixed linear model with genetic groups was fitted to the data to estimate race merit and the variance and covariances between traits, ages, and sites.

Results

Race effects for growth were small and only significant at the older site. In contrast, races differ significantly for height to phase change and density. The Colombian landrace and South and NE Tasmania races changed leaf type at a higher tree height. King Island and Recherche Bay had low density values, whereas the Colombian landrace had the highest. Heritability was moderate for growth traits (between 0.09 and 0.40), high for height to phase change (between 0.42 and 0.69), and moderate for wood density (0.28). The genetic correlation between growth and height to phase change was in general positive, but variable across sites. There was no correlation between growth and density. Despite large differences in growth between trials, pairwise genetic correlations suggest that genotype-by-site interaction is negligible. However, there was a poor correspondence between first-generation (ex-native stands) and second-generation families (ex-multi-provenance progeny trial). This suggests that breeding value estimates based on native open-pollinated material are unreliable.

Conclusion

Results confirm negligible race differences for growth, but not for wood density. Future breeding efforts should include the various genetic backgrounds including the Colombian landrace which constitutes a very important source, especially because of its high density. Despite a low Genotype by Environment interaction, the poor genetic correspondence between the first- and second-generation trials indicates that selections for growth should be based on information coming mainly from the advanced-generation trials.  相似文献   

11.

Context

Stem straightness is an important trait for growers and processors of Eucalyptus globulus logs for solid-wood products.

Aims

The aims of the study were to determine the extent of genetic variation in stem deviation from straightness in E. globulus and assess the utility of a six-point subjective scoring method as a selection criterion for stem straightness.

Methods

Two E. globulus progeny trials, grown under solid-wood product regimes, were studied. At age 9 years (post-thinning), stem straightness was measured using both image analysis and a six-point subjective scale. Diameter at breast height (DBH; 1.3 m) was measured at both age 5 (pre-thinning) and age 9 years.

Results

Significant additive genetic variation was observed. Strong, positive and significant additive genetic correlations were observed between the stem straightness assessment methods and between DBH at ages 5 and 9 years. Significant positive genetic correlations were shown between subjectively scored stem straightness and DBH at both ages 5 and 9 years.

Conclusion

The six-point subjective scoring method is a cost-effective selection criterion for stem straightness in E. globulus. The image measurement technique may be applied where objective estimates of stem straightness are required, for training purposes and to verify subjective scores.  相似文献   

12.

Aims

This study aims to evaluate the effects of wood ash application on nutrient dynamics and soil properties of an acidic forest soil (Arenosol).

Methods

Treatments were loose and pelleted ash application (11?Mg?ha?1), alone or together with N fertiliser, and control treatment in a lysimeter experiment. Nutrient leaching was followed during a 2-year period and soil chemical and biological properties were evaluated at the end of the experiment.

Results

Wood ash increased leaching of total N, NH 4 + -N, base cations and P, mainly during the first months, the effect being more pronounced for the loose formulation. At the end of the study period, a positive effect on soil nutrient availability and soil acidity reduction was seen. The application of loose and pelleted ash alone decreased N leaching and increased N microbial biomass at the end of the experiment. The C dynamics was weakly affected.

Conclusion

Wood ash can be used to improve nutrient availability and balance nutrient exported by tree harvesting in acid forest soils, the effects at short-term being stronger for loose than for pelleted ash. However, their application should be carried out when vegetation is established to minimise nutrient losses at short-term and reduce the potential risk for water bodies. In N-limited soils, wood ash should be applied with N fertilisers to counteract N immobilisation.  相似文献   

13.

Context

Avoidance or control of epicormic shoots is among the major silvicultural challenges for the production of high-quality oak timber. In northern Europe, contemporary oak silviculture aims to produce valuable timber on a relatively short rotation, applying early, heavy thinning combined with artificial pruning.

Aims

The aim of this study was to analyse the effects of pruning and stand density on the production of new epicormic shoots on young trees of pedunculate oak (Quercus robur L.).

Methods

The study was based on two field experiments in even-aged stands of pedunculate oak subjected to different thinning practices and early selection of potential future crop trees. From ages 13 to 15 years, stem density was reduced to 300 trees ha?1, 1,000 ha?1 or stands remained unthinned. Pruning was conducted on selected trees at ages 22–24 years. At that age, the stem density in unthinned control plots ranged from 2,500 to 3,100 ha?1. All treatments were replicated twice within each experiment.

Results

Pruning led to an overall increase in the total production of new epicormic shoots. More epicormic shoots were produced in the lower part of the stem (0–3 m in height) than in the upper part (3–6 m). The number of new epicormic shoots increased with increasing stand density.

Conclusion

Early, heavy thinning combined with high pruning at regular intervals may help shorten the rotation length for pedunculate oak without further reduction in wood quality than that which is caused by wider annual growth rings.  相似文献   

14.

? Context

While historical increases in forest growth have been largely documented, investigations on historical wood density changes remain anecdotic. They suggest possible density decreases in softwoods and ring-porous hardwoods, but are lacking for diffuse-porous hardwoods.

? Aims

To evaluate the historical change in mean ring density of common beech, in a regional context where a ring-porous hardwood and a softwood have been studied, and assess the additional effect of past historical increases in radial growth (+50 % over 100 years), resulting from the existence of a positive ring size–density relationship in broadleaved species.

? Methods

Seventy-four trees in 28 stands were sampled in Northeastern France to accurately separate developmental stage and historical signals in ring attributes. First, the historical change in mean ring density at 1.30 m (X-ray microdensitometry) was estimated statistically, at constant developmental stage and ring width. The effect of past growth increases was then added to assess the net historical change in wood density.

? Results

A progressive centennial decrease in mean ring density of ?55 kg?m?3 (?7.5 %) was identified (?10 % following the most recent decline). The centennial growth increase induced a maximum +25 kg?m?3 increase in mean ring density, whose net variation thus remained negative (?30 kg?m?3).

? Conclusions

This finding of a moderate but significant decrease in wood density that exceeds the effect of the positive growth change extends earlier reports obtained on other wood patterns in a same regional context and elsewhere. Despite their origin not being understood, such decreases hence form an issue for forest carbon accounting.  相似文献   

15.

Key message

Growth and wood chemical properties are important pulpwood traits. Their narrow-sense heritability ranged from 0.03 to 0.49 in Eucalyptus urophylla × E. tereticornis hybrids, indicating low to moderate levels of genetic control. Genetic correlations were mostly favorable for simultaneous improvement on growth and wood traits. Additive and non-additive genetic effects should be considered in making a hybrid breeding strategy.

Context

Eucalypt hybrids are widely planted for pulpwood production purposes. Genetic variations and correlations for growth and wood chemical traits remain to be explored in Eucalyptus interspecific hybrids.

Aims

Our objectives were to clarify the heritability of growth and wood chemical traits and determine the genetic correlations between traits and between trials in E. urophylla × E. tereticornis hybrids.

Methods

Two trials of 59 E. urophylla × E. tereticornis hybrids derived from an incomplete factorial mating design were investigated at age 10 for growth (height and diameter) and wood chemical properties (basic density, cellulose content, hemi-cellulose content, lignin content, and syringyl-to-guaiacyl ratio). Mixed linear models were used to estimate genetic parameters.

Results

Narrow-sense heritability estimates were 0.13?0.22 in growth and 0.03?0.49 in wood traits, indicating low to moderate levels of additive genetic control. Genetic correlations were mostly positively significant for growth with basic density and cellulose content but negatively significant with hemi-cellulose and lignin contents, being favourablefavorable for pulpwood breeding purpose. Type-B correlations between sites were significant for all the traits except diameter and lignin content.

Conclusion

Hybrid superiority warrants the breeding efforts. An appropriate breeding strategy should be able to capture both additive and non-additive genetic effects.
  相似文献   

16.

Aims

The aim of this study was to test the hypothesis that heavy predation of fallen fruits reduces the predation of neighboring post-dispersed seeds.

Methods

We performed a seed/fruit removal cafeteria experiment to assess the effects of seed species, of fruit species, and of density of fallen fruits on seed predation by wood mice (Apodemus sp.). In total, 6,930 seeds and 9,660 fruits from seven species were used. A generalized linear mixed model procedure was used to test these effects.

Results

Seed predation was significantly affected by the species of both seeds and neighboring fruits, but not by their density. Multiple comparison tests showed that most of treatments tested had no significant effects of species and density of neighboring fruits on seed removal. However, seed removal was significantly lower in some treatments with fruits than in the treatments without fruits for one of the seven seed species tested. Seed removal of two species tested was significantly differed among some treatments with different fruit species.

Conclusion

Most of the results in this study do not support our overall hypothesis. However, a part of results supported our hypothesis, suggesting that seed removal is affected by complex interactions between the preference of wood mice for fruits and fruiting environment.  相似文献   

17.

Key message

Pilodyn and acoustic velocity measurements on standing trees, used for predicting density and stiffness, can be good genetic selection tools for black spruce. Genetic parameters and selection efficiency were conserved in two breeding zones with contrasted bioclimatic conditions.

Context

Given the recent progress made in the black spruce genetic improvement program, the integration of juvenile wood mechanical properties as selection criteria is increasingly relevant.

Aims

This study aims to estimate the genetic parameters of in situ wood density and modulus of elasticity (MoE) measurements and to verify the efficiency of various measuring methods used for large-scale selection of black spruce based on wood quality.

Methods

Height, diameter, wood density, and some indirect measures of density (penetration and drilling resistance) and MoE (acoustical velocity and Pilodyn) were estimated on 2400 24-year-old trees of 120 open-pollinated families in progeny trials located in the continuous boreal or mixed forest subzones.

Results

Heritability of growth, density, and indirect density measurements varied from low to moderate and was moderate for acoustical velocity in both vegetation subzones. Expected genetic gains for wood properties based on in situ methods were higher for MoE proxy estimation combining Pilodyn and acoustic velocity.

Conclusion

Acoustic velocity is a good predictor of MoE. It is virtually unaffected by the environment and can be used on a large scale in the same manner as the Pilodyn for density. Using a proxy estimation that combines both methods helps optimize genetic gain for MoE.
  相似文献   

18.

Context

Climate change is expected to increase forest vulnerability through disturbances such as windstorms and droughts. Forest managers are therefore investigating strategies to increase forest resistance and resilience, especially by promoting uneven-aged and mixed forests through group selection, and by reducing stand stocking and large trees proportion. However, there is little information on the long-term impacts of these two practices.

Aims

The objectives of this study were (1) to develop an original silviculture algorithm designed for uneven-aged management and (2) to use it to assess the effects of the above-mentioned management methods in long-term simulations.

Methods

We simulated individual and group selection techniques in order to study the effects of group size, harvesting intensity and their interactions on wood production, stand heterogeneity, and regeneration in mountain spruce–fir forests. We used the spatially explicit individual-based forest model Samsara2 to simulate forest dynamics.

Results

Our simulation results confirmed the positive effect of group selection practices on structure diversity and regeneration but not on spruce maintenance. Increasing harvesting intensity enabled forest destocking but decreased structure diversity and led to non-sustained yields for the most intensive scenarios.

Conclusion

As adaptation measure, we thus recommend moderate group selection harvesting creating 500 m2 gaps.  相似文献   

19.
  • ? Decay resistance of larch (Larix sp.) to fungi was evaluated on heartwood samples belonging to 3 species (L. decidua, L. kaempferi and their hybrid), 3 races of European larch (polonica, sudetica and alpine), 13 wood lots (populations) and 313 trees.
  • ? Larch wood appeared, on average, as moderately durable although a high variability was observed. At the sample level as well as at the mean individual tree level, durability ranged from class 1 to 5 according to EN 350-1 standard. At the population level, larch wood varied from ‘durable’ to ‘slightly durable’. Genetics played a major role in decay resistance at the species, provenance and tree levels. Environmental factors such as the position of heartwood samples and the age of trees were also identified as a source of variability.
  • ? The most durable wood was not necessarily from old native alpine stands of European larch: some young larches from faster growing lowland origins also produced durable wood.
  • ? Genetic improvement of larch wood durability appeared therefore likely by the selection of the best populations for decay resistance as well as from the selection of individuals.
  •   相似文献   

    20.

    Context

    Current production from natural forests will not satisfy future world demand for timber and fuel wood, and new land management options are required.

    Aims

    We explore an innovative production system that combines the production of short rotation coppice in wide alleys with the production of high-value trees on narrow strips of land; it is an alternative form of alley cropping which we propose to call ‘alley coppice’. The aim is to describe this alley coppice system and to illustrate its potential for producing two diverse products, namely high-value timber and energy wood on the same land unit.

    Methods

    Based on a comprehensive literature review, we compare the advantages and disadvantages of the alley coppice system and contrast the features with well-known existing or past systems of biomass and wood production.

    Results

    We describe and discuss the basic aspects of alley coppice, its design and dynamics, the processes of competition and facilitation, issues of ecology, and areas that are open for future research.

    Conclusion

    Based on existing knowledge, a solid foundation for the implementation of alley coppice on suitable land is presented, and the high potential of this system could be shown.  相似文献   

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