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The structure of forest stands changes through developmental phases. This study is carried out in the unmanaged, oriental beech (Fagus orientalis Lipsky) stands in the north of Iran. The aim of this research was to quantify structural characteristics of stands in the stem exclusion phase using common structural indices, which include mingling, tree–tree distance, stem diameter, and tree height differentiation. According to our measurements from three stands, naturally regenerated stands tend to be mixed in species composition have slightly heterogeneous diameter distributions and uniform tree height. The average distance between trees was 3.3 m. Stocking volume of the stands had an average of 540 m3 ha?1 and 412 stem ha?1. Dead wood volume was 24 m3 ha?1, and as a standing volume, the most frequent species in dead wood pool was oriental beech (F. orientalis) (48 %). The common form of dead trees was snag (41 %). The mean value of mingling and tree-to-tree interval indices revealed that beech was mixed intensively with hornbeam and appears to be a more successful competitor for space and light compared with hornbeam; moreover, we found relatively high evidence of inter-species competition in this phase. A better understanding of stand characteristics in the stem exclusion phase as a critical part of the natural dynamics of forest ecosystems could facilitate predictions about the future changes within the stand.  相似文献   

3.
Mortality analysis of tree populations is widely applied to forest science studies. When mortality analysis is applied to forest inventories, the use of the variable of tree age is not common due to the difficulty of measuring age data censored in successive observations. The purpose of this study is to apply age-based mortality analysis, i.e., survival analysis, to the individual tree populations in a natural forest. The study site was the secondary natural stand of 0.26 ha dominated by fir (Abies firma), hemlock (Tsuga sieboldii), and oak (Quercus serrata) in the Boso peninsula, Japan. First, we measured the ages of the trees with diameter at breast height greater than or equal to 5 cm using a RESISTOGRAPH. Then, tree mortality probabilities of both non-parametric Kaplan–Meier estimates and parametric probability distributions of Gamma and Weibull were estimated by applying survival analysis techniques to the tree age data. The results implied that the survival analyses could be implemented not only by the non-parametric estimates in any cases but also by the common parametric distributions of Gamma and Weibull in cases in which the tree mortality probability distribution had a monotonously decreasing shape as observed in the immature natural stand in the study site.  相似文献   

4.
In the analysis of old-growth forest dynamics, the continuous process of tree aging and forest structural change is split up into several distinct forest development stages. The criteria for distinguishing the stages vary among the different approaches. In most of them, vertical canopy heterogeneity is only coarsely addressed and horizontal forest structure is quantified at spatial scales far exceeding the size of conventional forest inventory plots. In order to describe and analyze the complex mosaic structure of temperate old-growth forests with objective and quantitative measures in the context of forest inventories, we propose the Development Stage Index I DS . It employs two easily measured stand structural parameters (stem density and basal area) for quantifying the abundance of trees in three conventionally recognized tree diameter classes (premature ?<?40 cm; mature 40–70 cm; and over-mature?≥?70 cm) in plots of 500 m2 size, systematically distributed in the forest. This allows quantifying the spatial extension of the Initial, Optimum and Terminal stages of forest development at plot, stand and landscape levels. Based on thorough stand structural analyses in three virgin beech (Fagus sylvatica) forests in Slovakia, we demonstrate that I DS is a promising tool for (1) quantifying the proportion of the three stages on different scales, (2) visualizing the complex mixing of stages, and (3) analyzing dynamic changes in old-growth forest structure. We conclude that the Development Stage Index has the potential to improve the empirical foundation of forest dynamics research and to allow this discipline to proceed to more rigorous hypothesis testing.  相似文献   

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本文以杉木人工林和落叶松天然林标准地数据为材料,应用D=aN~bP~cS~dH~e数学模型进行计算分析,并从理论上证明,当同一树种的树高一定时,疏密度和株数相同,各地位指数(级)林分平均直径一致。为森林的抚育间伐,林分直径生长的预测、预报,确定林分工艺成熟龄及森林调查设计等提供了理论依据和重要参数。  相似文献   

7.
Over the last few decades, abandonment of traditional management practices in Spain has led to widespread stand densification and has favoured the expansion of some forest species that previously exhibited more restricted ranges. Spanish juniper (Juniperus thurifera L.) woodlands are experiencing this phenomenon due to agricultural land abandonment and a decrease in the livestock pressure. Yet the main drivers underlying stand structure and dynamics at this novel scenario are poorly understood. In this study, we investigate the main biotic drivers of tree growth in a high-density stand of the dioecious J. thurifera at an early developmental stage (mean tree age of 32, ca. 50 years after land abandonment). Tree growth was measured by coring 299 individuals of different reproductive classes (male, female and non-reproductive). Neighbourhood models were used to assess the relative importance of tree size and neighbourhood competition on tree growth of each reproductive class in the study plot. We found that tree size had the strongest effect on tree growth, whereas the effect of intraspecific competition was negligible. We observed differences in growth patterns among reproductive classes along trunk diameter sizes. Thus, at smaller sizes, the three reproductive classes presented identical patterns of growth. However, at bigger sizes, females were the fastest growing individuals, followed by males and non-reproductive individuals. Overall, our results suggest that in young J. thurifera monospecific forests, where self-thinning processes may have not undergone yet, tree size and the reproductive class could play a relatively more important role than competition as drivers of tree growth. These findings constitute new information which contributes to understanding growth dynamics at early developmental stages in this dioecious species. Furthermore, our results provide guidelines for silvicultural managing, suggesting that at these young juniper stands thinning would likely not translate into enhanced growth on remnant trees.  相似文献   

8.
Growth responses to thinning can vary with site quality and age, however, the direction of the response varies. An understanding of the mechanisms behind thinning responses could help forest managers optimise production as well as inform modellers and ecologists about the functioning of tree stands. Thinning was used to create a range of stand densities in eleven Eucalyptus plantation stands on seven sites in south-eastern Australia. Basal area periodic annual increment (PAI; cm2 year?1) of individual dominant and codominant trees was then related to competition, such that PAI = a + b ln(Competition). Competition was defined as the sum of the basal area of neighbouring trees within a radius of 6.5 m. The relative (%) and absolute (cm2 year?1) responses to competition were quantified using b, which was correlated with site quality and stand structure of unthinned stands. Stand structure was quantified using statistics or parameters that describe the diameter distribution for the given age, species and site, including skewness, the coefficient of variation and parameters of the Weibull probability distribution. Relative and absolute responses both increased with increasing site quality and in stands with more negatively skewed diameter distributions (higher proportion of larger trees) or lower coefficients of variation. Absolute thinning responses often increased with increasing size class, while relative thinning responses often decreased. Variables describing diameter distributions (size-class structure) were able to describe some of the variation in competition responses that site quality could not. This indicates why stands on similar site qualities but with different stand structures can have correspondingly different thinning responses. Stand structural variables may be more useful predictors of thinning responses in stands that experience large temporal changes in diameter distributions compared with site quality, which is a more static variable. Thus, knowledge about the development of diameter distributions might help to refine thinning regimes.  相似文献   

9.
天然林区小班森林资源数据的更新模型   总被引:22,自引:2,他引:20  
以吉林省汪清林业局为例,根据1997年森林经理调查的848块固定样地数据,与全林整体模型方法相结合,建立了适合于天然林区林业局(场)无人为干预小班森林资源数据更新的林分级生长模型组。该组模型包括林分密度指数,平均高,断面积,形高,郁闭度等林分测算因子的生长或变化模型。  相似文献   

10.
麻池背油松天然林林分生长结构的研究   总被引:7,自引:0,他引:7  
依据历年调查资料,对麻池背油松天然林的林分生长结构进行了研究.该林分的植物种类丰富;直径、树高、林龄等林分结构呈正态分布、近似正态分布和多峰状分布;近熟林的林分蓄积生长率为0.82%,材积生长量69年生时连年生长量仍大于平均生长量.由于林分密度过大,油松形状比值偏高,林木自然稀疏强烈,枯损率达32.7%,其中小径木占86.3%,病虫害感染枯死木占29.4%.油松天然更新不良,辽东栎不能成为更新树种,林下有青木杄、落叶松幼树分布.  相似文献   

11.
天然林分生长模型在小班数据更新中的应用   总被引:20,自引:0,他引:20  
当小班调查得到各组成树种的平均高,平均直径以及小班的年龄,每公顷株数,郁闭度,散生木蓄积等因子后,应用天然林分生长模型可以实现无人为干预小班数据的全面更新和连续预测。这些模型应用于小班数据更新时的特点是分别组成树种,以地位级指数评价立地质量,以预估间隔期代替年龄,更适用于天然混交异龄林小班数据的更新。通过吉林省汪清林业局108块复位样地的验证,表明天然林分生长模型的预估结果优于目前生产单位所用的生  相似文献   

12.
Sustainable forest management requires knowledge of forest structure and dynamics as well as an estimation of growing stock. The forest inventory provides the data for estimating stand variables. The measurement device MU2005-01738, patented by the Center for Forest Research (INIA-CIFOR), provides stereoscopic hemispherical images which can allow the 3D restoration of the stand around the sampling point. The aim of this study is to develop a methodology for forest stand mapping as well as tree diameter and height measurement along with volume estimation from the stereoscopic hemispherical images provided by the MU2005-01738. Using the MU2005-01738, Eucalyptus globulus Labill. plantations were sampled. Distance, diameter, height and volume were derived from the stereoscopic hemispherical images of 30 trees located at distances ranging from 0 to 15 m from the device. These variables were then compared with field measurements and the estimation errors analyzed. The (root-mean-squared error) RMSE was 0.23 m (8.95 %) for tree position and 1.51 cm (10.43 %) for diameter at breast height measurement at distances of less than 8 m. In the case of stem height and individual tree volume estimation, the RMSE was 2.59 m (23.05 %) and 0.025 m3 (17.94 %), respectively. The analysis of measurement errors indicated that the measurement precision decreases beyond 8 m from the device as well as for directions close to the baseline (the line between the optical centers of the two images), whereas the precision was highest for directions near to the line which is perpendicular to the base line. Future research should focus on improving measurement accuracy and possible applications in the field of forestry of the techniques presented in this study.  相似文献   

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The Bia owie a Primeval Forest, Poland, is the only forest area in temperate Europe where large tracts of natural forest communities have been conserved. The concept of silvatic mosaic [Oldeman, R.A.A., 1990. Forests: Elements of Silvology. Springer, Berlin] was applied in order to characterise the structure of stands representing oak–lime–hornbeam forest in both, the restricted area of Bia owie a National Park (BNP) and the commercial forests (CF), as well as to indicate a holistic approach towards protection and restoration of natural areas.Six developmental phases of stands were distinguished and defined: regeneration, young, pole, late pole, optimal, and terminal. Compositional and structural differences between the first five phases were also evaluated and visualised. Moreover, the considerable differences between analogous phases in BNP and CF were found. The uniform stand patches were mapped into two plots in BNP (25 ha each) and two in CF (25 and 20 ha), with a resolution of 10 m× 10 m pixels.The mosaic composition of natural communities indicates close-to-the equilibrium stage: the biostatic optimal phase occupies 33–40% of the area, while aggradational younger phases share nearly the same portion of the area as the senescent phase does. Although small, 0.01–0.04 ha eco-units dominate in BNP, there is a considerable portion of intermediate and large patches, indicating a complex character of factors responsible for the mosaic structure of natural stands in BNP. Both, composition and texture analyses indicate that the area of 25 ha is close to the minimum structure area specific for the communities studied.The comparison of natural and managed stands revealed large deficits of the optimal phase (only 19%) and a total absence of the terminal one in the managed area (20–35% in natural forest). Also the mosaic texture of CF plots was simplified because of standardisation of the management unit size. The modification of developmental phases, mosaic composition, and structure must lead to changes in the dynamics of the entire community. The present study demonstrates that the natural forest should be considered as a dynamic supersystem, covering a sufficient area to allow the developmental phases to perform in correct proportions. Because of differences in the durability of various phases, the natural cycle of forest development has an asynchronous and highly unpredictable character. Indeed, the applicability of multipurpose forestry rules as a tool of nature conservation has been questioned. An absolute ban on cutting in the remaining old-growth stands should be a basic rule for the restoration management.  相似文献   

15.
在杉木直径生长方程D=Dmt/(K+t)的基础上,进一步提出了杉木蓄积生长方程:Y=Ymt3/(K+t)3.蓄积方程和直径方程有一个共同的参数——林木生长特征系数K,这一特征系数K为杉木蓄积生长曲线的拐点.检验表明,蓄积Y的1/3次方与直径D基本保持线性相关,方程参数Ym(蓄积生长极值)和K与立地和密度因子的相关性符合和遵循林木生长基本规律和特点  相似文献   

16.
Heterogeneity of forest stand structure often results from repeated small disturbances, but structural variation also arises in a stand that has regenerated after a single large stand-replacing disturbance. We explored the structural variation within a subalpine Abies forest in Japan that regenerated after a large typhoon in 1959. In 2004, four 50 × 50 m plots were established at two sites in the regenerated forest. To characterize local stand structure within each plot, we determined the stem density, stand basal area, mean diameter at breast height (DBH), and coefficient of variation (CV) of DBH in 10 × 10 m subplots. We analyzed the spatial distribution pattern of the trees in each subplot using the L(t) function and categorized the distributions as clumped or non-clumped. The analysis revealed marked variation among subplots in the stand’s structural characteristics. Although the spatial distribution patterns of the trees in all plots were clumped, 11 of 50 subplots at one site and 39 of 50 subplots at the other site were non-clumped. Subplots with a clumped distribution pattern generally had a smaller basal area, smaller mean DBH, and greater CV of DBH than subplots with the same density but a non-clumped pattern. These results illustrated the spatial heterogeneity of forest structure that can arise in Abies forest that has experienced a large disturbance, probably because of the different densities and distribution of saplings surviving the disturbance and the different forest growth dynamics that result.  相似文献   

17.
Selective logging is the most widely employed method of commercial timber production in Asia, and its impact on forest structure, composition, and regeneration dynamics is considerable. However, the successional processes in forest communities after logging in semiarid mountains are poorly understood. To provide more information on these processes, we used data from tree rings, direct and indirect age determinations, and field measurements of stand structure to reconstruct the historical disturbance regime, stand development patterns, and successional processes in a natural Picea crassifolia forest community in the Qilian Mountains of northwestern China. The results showed that the density of P. crassifolia forest increased significantly after logging. The densities of second growth forests 30 and 70 years after logging disturbance had increased to 2874% and 294% of primary forest's density, respectively. Logging disturbance did not alter tree species composition of logged stands. However, the diversity of understory species changed significantly among the successional phases. Logging disturbance decreased the spatial heterogeneity of second growth forest. The spatial distributions of recruitment were affected by the location of the remaining trees. There was less recruitment near the remaining trees than near forest that had been cut. In addition, logging disturbance also induced a growth release for the trees on the sites sampled. Our results imply that the succession and regeneration of P. crassifolia forest may be improved if the remaining trees could be retained relative uniform distribution pattern, thinning or selective logging could be performed to height density, exotic shrubs could be removed or the shrubs cover could be reduced during the earlier successional stages.  相似文献   

18.
It has long been established that mature forests are mosaics of patches in different development phases but it has seldom explicitly been taken into account in ecological studies. We demonstrate here that these development phases, which are related to the population dynamics of trees, play an important role in the distribution of fauna based on observations on frugivorous birds. In an area close to the Calakmul Biosphere Reserve in Mexico, we studied the abundance of large forest bird species in relation to forest development phases, with a methodology that seems promising for ecological diagnosis and prognosis in forest management planning. Fine-scale forest mapping and bird counts were carried out in two block-transects of 40 m × 3000 m. Tree sampling in a sub-transect was used to generate population characteristics of trees. Large bird species preferred mature or senescent forest patches, whereas relatively young, growing forest patches were avoided. Important large tree species such as Manilkara zapota, Thouinia paucidentata, Guaiacum sanctum and Esenbeckia pentaphylla, characteristic of older forest patches, showed skewed size distributions indicating stress or overexploitation. The population of M. zapota, a key fruiting species that accounted for 26.5% of the total woody biomass, was most heavily affected by stress. A future collapse in the population of M. zapota, a decrease of the total area of older forest, and a decline in the abundance of large birds is likely if stress on the system continues at this level.  相似文献   

19.
近自然森林经营的基本理念是培育接近自然状态的森林,经营目标是恒续不断地发挥森林多种功能,基本方法与措施是单株作业和天然更新,参照对象是类似立地条件上的自然度近可能高的天然林,参照树种是天然林乔木层中的主、次要树种,目标树密度依据现有林分结构和林木生长发育情况而确定,确定目标直径的原则是"利益最大化",逐步疏开林冠,补植特殊目标树,将促使人工纯林稳步进入天然更新程序。  相似文献   

20.
Additional information concerning the quality of growing stock in forests has been obtained for the first time in Baden-Württemberg and Rheinland-Pfalz during the course of Germany’s second national forest inventory (BWI 2; conducted during the period 2001–2002). In this article, the quality assessment—called stem quality rating method—is described with a special focus on its potential to provide the basis for a more detailed investigation of the growing stock’s quality distribution. As a main result, the article presents and illustrates a model-based quantification of single tree and stand/site variable effects on the quality distribution of Norway spruce. Single tree variables showing a significant effect are diameter at breast height (DBH), height–DBH-ratio (h/d-value), age and distance to forest edge. Additional stand/site variables which have a significant effect are altitude, terrain slope, stand type and inventory team. Due to the ordinal type of the response variable, a categorical regression model is applied. Non-linear effects of predictor variables were detected and modeled by integration of smoothing spline terms. Validating model predictions with regard to expert knowledge in forestry led to the integration of simple constraints in the linear predictor, which controls whether category-specific effects are fitted or not. The resulting model could be described as a vector generalized additive non-proportional odds regression model. This improved insight into the determination of stem quality could be applied in optimization studies to derive optimal silvicultural treatments and in the setting up of management guidelines. Assuming constant relationships between predictor and response variables over time, the combined application with growth simulators allows for a prediction of future joint quality and size class assortment distributions. Finally, the model would allow for a sustainability control of stem quality over time if a consecutive inventory will be conducted during the course of the third German national forest inventory (BWI 3).  相似文献   

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