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We reviewed studies dealing with regeneration under variable conditions in boreal and hemiboreal forests as affected by different microsite types by tree species functional groups. Generally, the importance of storm-induced microsites for regeneration dynamics in boreal forests depends on several factors: (1) distribution and type of microsites (generated by storm characteristics and stand conditions); (2) viable seed supply (stand history, species dispersal traits and status of surviving trees) and their species’ life history strategy; (3) climatic and site conditions (pre-storm conditions and storm-induced changes); and (4) delayed storm effects, such as retarded falling of trees, favoured vegetation growth, etc. Studies acknowledging the significance of microsites were mostly related to intermediate or severe events, causing sufficient changes in resource levels and growth conditions, and influencing extrinsic factors such as frost heaving, erosion and browsing. Also, the dispersal traits of available tree species, including sprouting and response of surviving trees, such as canopy expansion, should be considered in evaluating microsite importance in individual cases. In intermediate to severe windstorm events, pioneer species are generally profiting most from the additional offer in microsites, requiring bare mineral soil and elevated locations for their establishment and growth. Under gap dynamics, shade-tolerant species benefit from dead wood and elevated locations as these offer safe sites in stands with abundant understorey vegetation.  相似文献   
2.
  • ? This article synthesizes and reviews the available information on the effects of forestry practices on the occurrence of biotic and abiotic hazards, as well as on stand susceptibility to these damaging agents, concentrating on mammal herbivores, pest insects, pathogenic fungi, wind and fire.
  • ? The management operations examined are site selection, site preparation, stand composition, regeneration method, cleaning and weed control, thinning and pruning, and harvesting. For each of these operations we have examined how they influence the occurrence of biotic and abiotic damaging agents, the susceptibility of European forests, and describe the ecological processes that may explain these influences.
  • ? Overall, we find that the silvicultural operations that have the largest influence on both biotic and abiotic risks to European forest stands are closely related to species composition and the structure of the overstorey. Four main processes that drive the causal relationships between stand management and susceptibility have been identified: effect on local microclimate, provision of fuel and resources to biotic and abiotic hazards, enhancement of biological control by natural enemies and changes in individual tree physiology and development.
  • ? The review demonstrates an opportunity to develop silvicultural methods that achieve forest management objectives at the same time as minimising biotic and abiotic risks.
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    3.

    The anticipated increase in extreme disturbance events due to climate change is likely to expose Norway spruce (Picea abies (L.) Karst.) dominated forests in northern Europe to new conditions. Empirical data on the resilience of such natural (unmanaged) forests to disturbance and the long-term patterns of regeneration in its aftermath are currently scarce. We performed a quantitative assessment of natural forest stands in north–western Latvia to identify and characterise the patterns of stand structure 44 years after a stand-replacing disturbance and investigated the effects of legacies on regeneration. The spatial distribution of tree species and their dimensions were assessed in 71 circular sample plots (500 m2 each) in natural forest areas, where Norway spruce dominated prior to the windthrow and salvage logging was not carried out. Spatial indices (species mingling index, size differentiation index, and aggregation index) were used to characterise stand structure and diversity. The different initial states (age and coverage of surviving trees) of stands affected eventual tree species dominance, size differentiation, degree of mingling and aggregation. Our results demonstrate a close relationship between disturbance legacies and spatial indices. The pre-storm understory and canopy survivors decreased species mingling, whereas survivors increased size differentiation. The size differentiation increased also with a higher degree of species mingling. Leaving differential post-storm legacies untouched promotes a higher structural and species diversity and therefore supports the management approach of preserving canopy survivors.

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    4.
    Natural regeneration of windthrow areas is an important issue when planning forestry measures after forest disturbances. Seedling recruitment was investigated in storm-damaged hemiboreal mixed forests in eastern Estonia. The establishment and growth of seedlings from natural regeneration was registered for tree species in soil pits and in mounds of uprooted trees in stands that were either heavily or moderately damaged. Seedling growth is expected to be better in large but shallow soil pits created by uprooted Norway spruce [Picea abies (L.) Karst.] and poorer in small but deep pits created by the hardwoods in the area, silver birch (Betula pendula Roth.) and European aspen (Populus tremula L.). The most abundant regenerating species was birch. Pits hosted larger seedling numbers than mounds, due to soil instability in mounds. Rowan (Sorbus aucuparia L.) showed significantly faster growth than the other seedling species. Norway spruce pits were preferred to pits of other species by both birch and spruce seedlings. Black alder [Alnus glutinosa (L.) J. Gaertn.] did not show a preference for pits of a certain species of uprooted tree. Both spruce and rowan preferred hardwood mounds over spruce mounds. Storm severity also affected species composition: birch predominantly occurred on pits and mounds in heavily disturbed areas, while spruce was more abundant in the moderately damaged areas. The effects of advance regeneration and surrounding stands on seedling microsite preferences should be considered in future research and subsequent management recommendations.  相似文献   
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