State Key Laboratory of Tea Plant Biology and Utilization, School of Tea and Food Science & Technology, Anhui Agricultural University, Hefei, China;2. School of Resource & Environment of Anhui Agricultural University, Key Laboratory of Agri-food Safety of Anhui Province, Hefei, China;3. State Key Laboratory of Tea Plant Biology and Utilization, School of Tea and Food Science & Technology, Anhui Agricultural University, Hefei, China;4. Department of Molecular Biosciences and Bioengineering, University of Hawaii at Manoa, Honolulu, HI, USA